If ‘O ^ P’ means ‘O is the brother of P’, ‘O * P’ means ‘O is the mother of…

If

‘O ^ P’ means ‘O is the brother of P’,

‘O * P’ means ‘O is the mother of P’,

'O + P’ means ‘O is the husband of P’,

‘O @ P’ means ‘O is the sister of P’,

‘O = P’ means ‘O is the son of P’, and

‘O %P’ means ‘O is the father of P’,

then how is S related to M the following expression?

S = G %H @ E + F * M