If ‘Z @ S’ means ‘Z is the father of S’, ‘Z * S’ means ‘Z is the brother of…

If ‘Z @ S’ means ‘Z is the father of S’, ‘Z * S’ means ‘Z is the brother of S’, 'Z × S’ means ‘Z is the father’s sister of S’, ‘Z # S’ means ‘Z is the sister of S’, ‘Z $ S’ means ‘Z is the daughter of S’, and ‘Z = S’ means ‘Z is the mother of S’, then how is P related to U in the following expression? P × Q$ R@ S * T $ U